Hi
I did analyze your G-Code and isolated part of it which is probably is causing issues.
The model.
here are related G-Code with analysis
So as I can see by numbers - your G-Code is overloading your extruder on retract and unretract.
If you would also account for Pressure_advance - then your real rates will become even higher, because when unretract is happening PA additionally is trying to create huge pressure in the nozzle so it could extrude at that high rate and your stepper-motor is just skipping (it can’t handle such high speeds).
Try to decrease retract/un-retract speeds to 20-30mm/s or 1200-1800mm/m
this will reduce the rate to 48-72mm^3/s and then after the re-slicing - re-check G-Code if it’s actually reduced.